Science and maths education

Why are formula triangles bad? Education in Chemistry,  June 2015 available online

What is peer review? Classroom resources. Education in Chemistry, June 2018, available online

Nine tips for scaffolding multi-step calculations. Education in Chemistry, October 2018, available online

Antibiotics: solving an evolving problem, Classroom resources. Education in Chemistry, November 2018, available online


Maths in science

J A Koenig and N Pike (2013). Perspectives from the UK and the US on integrating
mathematics into the teaching and learning of the biological sciences in higher education. Available online

J A Koenig (2011). A survey of the mathematics landscape within bioscience undergraduate and postgraduate UK higher education. UK Centre for Bioscience, now available here

J. A. Koenig (2011) The mathematical education of pharmacologists through school and undergraduate study. “Pharmacology Matters” magazine of the British Pharmacological Society.

J A Koenig (2007) It’s not rocket science. Higher Education Academy Centre for Bioscience Bulletin, Volume 21, Summer 2007

J A Koenig (2006) Essential maths for medics and vets. CETL-MSOR Conference 2006 Proceedings available online at



J A Koenig (2010) Drug Receptor Interactions: A Guide to Calculations.

J A Koenig (2010) A Guide to Calculations in Pharmacokinetics.


J A Koenig (2004). Signal reception: G protein-coupled receptors. In “Molecular Biology of the Neuron” Second Edition. Ed R W Davies and B J Morris, Oxford University Press.

J A Koenig (2004) Assessment of receptor internalization and recycling. Methods Mol Biol. 259, 249-73.

J Gonzalez-Maeso, A Wise, A Green and J A Koenig (2003). Agonist-induced desensitization and endocytosis of heterodimeric GABAB receptors in CHO-K1 cells. Eur J Pharmacol., 481, 15-23.

K S M Smalley, J A Koenig, W Feniuk and P P A Humphrey (2001). Ligand internalization and recycling by human recombinant somatostatin type 4 (h sst4) receptors expressed in CHO-K1 cells. Brit. J Pharmacol, 132, 1102−1110.

J A Koenig, R Kaur, I Dodgeon, J M Edwardson and P P A Humphrey, (1998). Fates of endocytosed somatostatin sst2 receptors and associated agonists. Biochem. J. 336, 291–298.

J A Koenig (1998). Radioligand Binding in Intact Cells. In “Receptor Binding Techniques” Ed M Keen. Humana Press

J A Koenig (1998). G protein Coupled Receptors. In “Molecular Biology of the Neuron” Edited by R. Wayne Davies and Brian J. Morris, Oxford University Press.

P G Szekeres, J A Koenig and J M Edwardson (1998). The Relationship between Agonist Intrinsic Activity and the Rate of Endocytosis of Muscarinic Receptors in a Human Neuroblastoma Cell Line. Mol Pharmacol., 53, 759-765.

P G Szekeres, J A Koenig and J M Edwardson (1998). Involvement of receptor cycling and receptor reserve in resensitization of muscarinic responses in SH-SY5Y human neuroblastoma cells. J Neurochem. 70,1694-703.

J A Koenig and J M Edwardson (1997). Endocytosis and recycling of G protein-coupled receptors. Trends in Pharmacological Sciences, 18, 276-287.

J A Koenig, J M Edwardson and P P A Humphrey, (1997). Brit J Pharmacol,120, 45-51.
Somatostatin receptors in Neuro2A neuroblastoma cells: operational characteristics.

J A Koenig, J M Edwardson and P P A Humphrey, (1997). Brit J Pharmacol,120, 52−59.
Somatostatin receptors in Neuro2A neuroblastoma cells: ligand internalization

J A Koenig and J M Edwardson, (1996). Intracellular trafficking of muscarinic actylcholine receptors: importance of subtype and cell type. Mol. Pharmacol., 49, 351-359.

J A Koenig and J M Edwardson, (1994). Kinetic Analysis of the Trafficking of Muscarinic
Acetylcholine Receptors between the Plasma Membrane and Intracellular Compartments. J Biol Chem, 269, 17174 – 82.

JA Koenig and J M Edwardson, (1994). Routes of delivery of muscarinic acetylcholine receptors to the plasma membrane in NG108-15 cells. Br J Pharmacol. 111, 1023-8

J A Koenig and I L Martin, (1992) Effect of free fatty acids on GABAA receptor ligand binding. Biochem Pharmacol. 44,11-5.

M E McManus, I Stupans, W Burgess,  J A Koenig, P M Hall and D J Birkett (1987). Flavin-containing monooxygenase activity in human liver microsomes. Drug Metabolism and Disposition, 15(2), 256-261.